JACKSON -- Biologists have confirmed that a female grizzly bear killed by a hunter last month in northwest Wyoming was from a well-known Grand Teton National Park bear family.

Park officials say genetic testing confirmed the dead grizzly was one of three offspring of a bear known as number 399.

Bear 399 raised her three cubs by the roadside near Jackson Lake Lodge in 2006-08, gaining some measure of fame among locals and tourists.

Authorities say Stephen Westmoreland, of Teton Village, shot one of its cubs on Sept. 19 from 40 yards away while he was helping carry a deer his hunting partner shot in Bridger-Teton National Forest.

Authorities have charged Westmoreland with taking a grizzly bear without a license after investigators said the shooting was not done in self defense.

Westmoreland's attorney, David DeFazio, of Jackson, was out of the state and not available for comment Friday. There is no Internet telephone listing for Westmoreland.

According to a Wyoming Game and Fish report, Westmoreland was walking along the trail in a 3-acre meadow when the bear appeared, scavenging a moose carcass left behind by another hunter.

Westmoreland was armed with a rifle and a pistol but was not carrying bear pepper spray. He shot the bear with the rifle from about 40 yards as the bear dropped to all fours and faced him, according to Game and Fish. The bear was shot first through the chest and then in the mid-body area on its left side. The bear died five feet from the moose carcass, according to the report.

Westmoreland reported the incident himself, saying he feared the bear would attack him because he was covered in animal blood after gutting a deer.

Grand Teton senior wildlife biologist Steve Cain said the 3-year-old female bear was habituated to people but had never become conditioned to human food.

"(She) had not been involved in any conflicts," he said. "The science indicates that bears that are habituated to people are less likely to act aggressively toward people."